A monograph and re-classification of the previous genus Limosina Macquart (Diptera, Sphaeroceridae) of Europe. Part II.

Abstract

The European species belonging to the previous genus Limosina Macquart (sensu Hackmann 1969a) are monographed. This taxon was found to be polyphyletic and is divided using cladistic classification into 17 genera with a total of 81 European species. 11 genera and 11 subgenera are discribed as new: Gigalimosina gen. nov., Herniosina gen. nov., Terrilimosina gen. nov., Minilimosina gen. nov., Minilimosina (Svarciella) subgen. nov., Minilimosina (Allolimosina) subgen. nov., Xenolimosina gen. nov., Paralimosina (Canarisina) subgen. nov., Spelobia (Eulimosina) subgen. nov., Spelobia (Bifronsina) subgen. nov., Pullimosina gen. nov., Pullimosina (Dahlimosina) subgen. nov., Spinilimosina gen. nov., Kimosina gen. nov., Kimosina (Kollimosina) subgen. nov., Kimosina (Alimosina) subgen. nov., Telomerina gen. nov., Opalimosina gen. nov., Opalimosina (Pappiella) subgen. nov., Opalimosina (Hakmanina) subgen. nov., Opalimosina (Dentilimosina) subgen. nov., and Rudolfia gen. nov. Other included (Limosina Maquart, 1835; Apteromyia Vimmer, 1929 gen. restit.; Paralimosina Papp, 1973; Spelobia Spuler, 1924 nom. restit., stat. nov.; Chaetopodella Duba, 1920; Halidayina Duba 1918) are rediscribed. Descriptions of 14 new species are given (Minilimosina 4, Spelobia 6, Kimosina 1, Telomeina 3), other are fully rediscribed. 4 new synonyms are established; numerous species (besides those included in the Monograph) are put in new combinations. All available type material of European species (including synonyms) was revised, number of lectotypes designated and in this way the status of particular species fixed. A description of each species is supplemented by numerous illustrations, data on the morphology of preimaginal stages, biology, distribution and by discussion of relationships. Identification keys to all included taxa are given. The general part includes a summary of basic data dealing with history of investigations, morphology, biology, zoogeography and phylogeny (cladistic classification) of the groups under study.

Nomenclatural Acts

claviventris (Strobl, 1909) (Apteromyia), comb. no. hitherto Limosina claviventris
flaviceps (Zetterstedt, 1847) (Gigalimosina), comb. n. hitherto Limosina flaviceps
bequaerti (Villeneuve, 1917) (Herniosina), comb. n. hitherto Leptocera (Limosina) bequaerti
horrida (Roháček, 1978) (Herniosina), comb. n. hitherto Limosina horrida
Alimosina Roháček, 1983 (Kimosina), sgen. n.
Collimosina Roháček, 1983 (Kimosina), sgen. n.
empirica (Hutton, 1901) (Kimosina (Alimosina)), comb. n. hitherto Borborus empiricus
spinosa (Collin, 1930) (Kimosina (Collimosina)), comb. n. hitherto Limosina spinosa
ciliata (Duda, 1918) (Kimosina (Kimosina)), comb. n. hitherto Limosina (Scotophilella) ciliata
glabrescens (Villeneuve, 1917) (Kimosina (Kimosina)), comb. n. hitherto Leptocera (Limosina) glabrescens
longisetosa (Dahl, 1909) (Kimosina (Kimosina)), comb. n. hitherto Limosina longisetosa
pappi Roháček, 1983 (Kimosina (Kimosina)), spec. n.
plumosula (Rondani, 1880) (Kimosina (Kimosina)), comb. n. hitherto Limosina plumosula
latigena Roháček, 1977 (Limosina), syn. n. of Paralimosina (Paralimosina) kaszabi Papp, 1973
Allolimosina Roháček, 1983 (Minilimosina), sgen. n.
Svarciella Roháček, 1983 (Minilimosina), sgen. n.
albinervis (Duda, 1918) (Minilimosina (Allolimosina)), comb. n. hitherto Limosina (Scatophilella) albinervis
alloneura (Richards, 1952) (Minilimosina (Allolimosina)), comb. n. hitherto Leptocedra (Limosina) alloneura
secundaria (Duda, 1918) (Minilimosina (Allolimosina)), comb. n. hitherto Limosina (Scatophilella) secundaria
fungicola (Haliday, 1836) (Minilimosina (Minilimosina)), comb. n. hitherto Limosina fungicola
gemella Roháček, 1983 (Minilimosina (Minilimosina)), spec. n.
parvula (Stenhammar, 1854) (Minilimosina (Minilimosina)), comb. n. hitherto Limosina parvula
tenera Roháček, 1983 (Minilimosina (Minilimosina)), spec. n.
trogeri Roháček, 1983 (Minilimosina (Minilimosina)), spec. n.
hackmani (Roháček, 1977) (Minilimosina (Svarciella)), comb. n. hitherto Limosina hackmani
ismayi Roháček, 1983 (Minilimosina (Svarciella)), spec. n.
splendens (Duda, 1928) (Minilimosina (Svarciella)), comb. n. hitherto Limosina (Scotophilella) splendens
v-atrum (Villeneuve, 1917) (Minilimosina (Svarciella)), comb. n. hitherto Leptocera (Limosina) v-atrum
vitripennis (Zetterstedt, 1847) (Minilimosina (Svarciella)), comb. n. hitherto Limosina vitripennis
Dentilimosina Roháček, 1983 (Opalimosina), sgen. n.
Hackmanina Roháček, 1983 (Opalimosina), sgen. n.
Pappiella Roháček, 1983 (Opalimosina), sgen. n.
denticulata (Duda, 1924) (Opalimosina (Dentilimosina)), comb. n. hitherto Limosina (Scotophilella)
czernyi (Duda, 1918) (Opalimosina (Hackmanina)), comb. n. hitherto Limosina (Scotophilella) czernyi
calcarifera (Roháček, 1975) (Opalimosina (Opalimosina)), comb. n. hitherto Limosina calcarifera
collini (Richards, 1929) (Opalimosina (Opalimosina)), comb. n. hitherto Leptocera collini
mirabilis (Collin, 1902) (Opalimosina (Opalimosina)), comb. n. hitherto Limosina mirabilis
simplex (Richards, 1929) (Opalimosina (Opalimosina)), comb. n. hitherto Leptocera grenstedi var. simplex
liliputana (Rondani, 1880) (Opalimosina (Pappiella)), comb. n. hitherto Limosina liliputana
Canarisina Roháček, 1983 (Paralimosina), sgen. n.
beckeri (Duda, 1918) (Paralimosina (Canarisina)), comb. n. hitherto Limosina (Scotophilella) beckeri
fucata (Rondani, 1880) (Paralimosina (Paralimosina)), comb. n. hitherto Limosina fucata
macedonica (Roháček, 1977) (Paralimosina (Paralimosina)), comb. n. hitherto Limosina macedonica
subcribrata (Roháček, 1977) (Paralimosina (Paralimosina)), comb. n. hitherto Limosina subcribata
trichopyga (Richards, 1952) (Paralimosina (Paralimosina)), comb. n. hitherto Leptocera (Limosina) trichopyga
Dahlimosina Roháček, 1983 (Pullimosina), sgen. n.
dahli (Duda, 1918) (Pullimosina (Dahlimosina)), comb. n. hitherto Limosina (Scotophilella) dahli
karelica (Papp, 1979) (Pullimosina (Dahlimosina)), comb. n. hitherto Limosina karelica
antennata (Duda, 1918) (Pullimosina (Pullimosina)), comb. n. hitherto Limosina (Scotophilella) antennata
heteroneura (Haliday, 1836) (Pullimosina (Pullimosina)), comb. n. hitherto Limosina heteroneura
meijerei (Duda, 1918) (Pullimosina (Pullimosina)), comb. n. hitherto Limosina (Scotophilella) meijerei
moesta (Villeneuve, 1918) (Pullimosina (Pullimosina)), comb. n. hitherto Limosina (Leptocera) moesta
pullula (Zetterstedt, 1847) (Pullimosina (Pullimosina)), comb. n. hitherto Limosina pullula
rozkosnyi (Roháček, 1975) (Rudolfia), comb. n. hitherto Limosina rozkosnyi
Bifronsina Roháček, 1983 (Spelobia), sgen. n.
Eulimosina Roháček, 1983 (Spelobia), sgen. n.
bifrons (Stenhammar, 1854) (Spelobia (Bifronsina)), comb. n. hitherto Limosina bifrons
ochripes (Meigen, 1830) (Spelobia (Eulimosina)), comb. n. hitherto Borborus ochripes
baezi (Papp, 1977) (Spelobia (Spelobia)), comb. n. hitherto Limosina baezi
belanica Roháček, 1983 (Spelobia (Spelobia)), spec. n.
cambrica (Richards, 1929) (Spelobia (Spelobia)), comb. n. hitherto Leptocera cambrica
clunipes (Meigen, 1830) (Spelobia (Spelobia)), comb. n. hitherto Borborus clunipes
czizeki (Duda, 1918) (Spelobia (Spelobia)), comb. n. hitherto Limosina (Scotophilella) czizeki
faeroensis (Deeming, 1966) (Spelobia (Spelobia)), comb. n. hitherto Leptocera (Limosina) faeroensis
ibrida Roháček, 1983 (Spelobia (Spelobia)), spec. n.
luteilabris (Rondani, 1880) (Spelobia (Spelobia)), comb. n. hitherto Limosina luteilabris
manicata (Richards, 1927) (Spelobia (Spelobia)), comb. n. hitherto Limosina manicata
nana (Rondani, 1880) (Spelobia (Spelobia)), comb. n. hitherto Limosina nana
palmata (Richards, 1927) (Spelobia (Spelobia)), comb. n. hitherto Limosina palmata
pappi Roháček, 1983 (Spelobia (Spelobia)), spec. n.
parapusio (Dahl, 1909) (Spelobia (Spelobia)), comb. n. hitherto Limosina parapusio
pseudonivalis (Dahl, 1909) (Spelobia (Spelobia)), comb. n. hitherto Limosina pseudonivalis
pseudosetaria (Duda, 1918) (Spelobia (Spelobia)), comb. n. hitherto Limosina (Scotophilella) pseudosetaria
quaesita Roháček, 1983 (Spelobia (Spelobia)), spec. n.
rufilabris (Stenhammar, 1854) (Spelobia (Spelobia)), comb. n. Limosina
simplicipes (Duda, 1925) (Spelobia (Spelobia)), comb. n. hitherto Limosina (Scotophilella) simplicipes
talis Roháček, 1983 (Spelobia (Spelobia)), spec. n.
talparum (Richards, 1927) (Spelobia (Spelobia)), comb. n. hitherto Limosina taplarum
ulla Roháček, 1983 (Spelobia (Spelobia)), spec. n.
villosa (Duda, 1918) (Spelobia (Spelobia)), comb. n. hitherto Limosina (Scotophilella) villosa
Aptilomyia Vimmer, 1929 (Sphaeroceridae), syn. n. of Apteromyia Vimmer, 1929
Gigalimosina Rohácek, 1983 (Sphaeroceridae), gen. n.
Herniosina Rohácek, 1983 (Sphaeroceridae), gen. n.
Kimosina Roháček, 1983 (Sphaeroceridae), gen. n.
Minilimosina Roháček, 1983 (Sphaeroceridae), gen. n.
Opalimosina Roháček, 1983 (Sphaeroceridae), gen. n.
Pullimosina Roháček, 1983 (Sphaeroceridae), gen. n.
Rudolfia Roháček, 1983 (Sphaeroceridae), gen. n.
Spelobia Spuler, 1924 (Sphaeroceridae), stat. n. hitherto subgenus of Leptocera
Spinilimosina Roháček, 1983 (Sphaeroceridae), gen. n.
Telomerina Roháček, 1983 (Sphaeroceridae), gen. n.
Terrilimosina Roháček, 1983 (Sphaeroceridae), gen. n.
Vseteckiella Vimmer, 1931 (Sphaeroceridae), syn. n. of Apteromyia Vimmer, 1929
Xenolimosina Roháček, 1983 (Sphaeroceridae), gen. n.
brevicostata (Duda, 1918) (Spinilimosina), comb. n. hitherto Limosina (Scotophilella) brevicostata
antonini Roháček, 1983 (Telomerina), spec. n.
eburnea Roháček, 1983 (Telomerina), spec. n.
flavipes (Meigen, 1930) (Telomerina), comb. n. hitherto Borborus flavipes
pseudoleucoptera (Duda, 1924) (Telomerina), comb. n. hitherto Limosina (Scotophilella) pseudoleucoptera
ursina Roháček, 1983 (Telomerina), spec. n.
racovitzai (Bezzi, 1911) (Terrilimosina), comb. n. hitherto Limosina racovitzai
schmitzi (Duda, 1918) (Terrilimosina), comb. n. hitherto Limosina (Scotophilella) schmitzi
sudetica (Roháček, 1975) (Terrilimosina), comb. n. hitherto Limosina sudetica
setaria (Villeneuve, 1918) (Xenolimosina), comb. n. hitherto Limosina (Leptocera) setaria