Taxonomy and historical biogeography of the genus Rhyacophila Pictet, 1834 on the Philippines (Insecta, Trichoptera).

Abstract

Nine species of the genus Rhyacophila occur on the Philippines. Five species are described as new: R. baldensis sp. n., R. banahawmontis sp. n., R. amoyanensis sp. n., R. pilifera sp. n., and R. cordilliera sp. n. The male genitalia and adults of four species are illustrated. The species belong to two species-groups, with the latter group established herein. The ranges of these groups in the Philippines are depicted in two maps. Phylogenetic relationships were analysed and are presented in three cladograms based on morphological characters of male genitalia. The species of the castanea-group immigrated into the Philippines via Borneo, whereas the species of the spinosellata-group are relict elements of an older Asian fauna. A dispersal scenario is proposed for both groups concerning the colonisation of the Philippines.

Keywords

Insecta, Trichoptera, Rhyacophila Pictet, new species, taxonomy, phylogeny, historical biogeography, dispersal scenario, Borneo, the Philippines.

Nomenclatural Acts

amoyanensis Mey, 2011 (Rhyacophila), spec.nov.
baldensis Mey, 2011 (Rhyacophila), spec.nov.
banahawmontis Mey, 2011 (Rhyacophila), spec.nov.
cordilliera Mey, 2011 (Rhyacophila), spec.nov.
pilifera Mey, 2011 (Rhyacophila), spec.nov.