On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae).

Abstract

Types and additional material of twelve genera, one of them new, of Lomechusini from the Palaearctic and Oriental regions are revised. In all, 70 species are (re-)described and/or illustrated, 51 of them new: Orphnebius bakeri Bernhauer, 1929; O. breviceps Cameron, 1946; O. opticus Cameron, 1946; O. bicuspis spec. nov. (Laos; India: Arunachal Pradesh); O. biformis spec. nov. (Indonesia: Sumatra); O. cernens spec. nov. (Laos); O. dilatatus spec. nov. (Laos); O. effeminatus spec. nov. (Malaysia); O. extensus spec. nov. (Laos); O. fodens spec. nov. (Malaysia: Sabah); O. fuscapicalis spec. nov. (Laos); O. fusicollis spec. nov. (Laos); O. grandicollis spec. nov. (Laos); O. integer spec. nov. (Laos); O. latitibialis spec. nov. (Laos); O. lunatus spec. nov. (Laos); O. nigrapicalis spec. nov. (Laos); O. reductus spec. nov. (Laos); O. retunsus spec. nov. (Laos); O. serratus spec. nov. (Laos); O. spinans spec. nov. (India: Arunachal Pradesh); Orphnebius (Deroleptus) falagrioides Bernhauer, 1929; O. (D.) laticeps Cameron, 1925; O. (D.) multimpressus Assing, 2015; O. (D.) niger (Cameron, 1939); O. (D.) siamensis Cameron, 1939; O. (D.) baccillatus spec. nov. (Laos); O. (D.) biimpressus spec. nov. (Indonesia: Sumatra); O. (D.) carinatus spec. nov. (Laos); O. (D.) cultellatus spec. nov. (Thailand, Laos); O. (D.) discrepans spec. nov. (China: Yunnan); O. (D.) dispar spec. nov. (India: Arunachal Pradesh); O. (D.) gracilior spec. nov. (India: Arunachal Pradesh); O. (D.) septemcuspis spec. nov. (Laos); O. (D.) sexcarinatus spec. nov. (Indonesia: Sumatra); O. (D.) spoliatus spec. nov. (Laos); O. (D.) tortus spec. nov. (India: Meghalaya); O. (D.) triapicalis spec. nov. (China: Sichuan); O. (D.) ulcerosus spec. nov. (Malaysia: Sabah); O. (D.) vates spec. nov. (Laos); Drusilla bifida spec. nov. (Thailand); Rabdotodrusilla pectinata spec. nov. (Thailand); Amaurodera meorum Pace, 1992; A. thailandensis Pace, 1986; A. arunica spec. nov. (Nepal); A. dentata spec. nov. (Thailand); A. disparicollis spec. nov. (Indonesia: Sumatra); A. fasciata spec. nov. (Thailand); A. gilvios spec. nov. (Thailand); A. parvoculata spec. nov. (Thailand); A. reticulata spec. nov. (Thailand); A. spinans spec. nov. (Indonesia: Sumatra); A. varicollis spec. nov. (Indonesia: Sumatra); Tetrabothrus borneensis Cameron, 1943; T. indicus Cameron, 1939; T. inflexus Assing, 2015; T. neoguineensis Pace, 2012; T. pubescens Bernhauer, 1915; T. breviatus spec. nov. (China: Sichuan); T. collucatus spec. nov. (Laos); T. nilgiricus spec. nov. (India: Tamil Nadu); T. punctiventris spec. nov. (Malaysia: Sabah); T. sulawesicus spec. nov. (Indonesia: Sulawesi Utara); Zyras (Zyras) gibbus Pace, 2010; Z. (Z.) illecebrosus Last, 1982; Z. (Z.) quasar Dvořak, 1996; Z. (Z.) porrectus spec. nov. (China: Sichuan); Z. (Z.) wunderlei spec. nov. (Indonesia: Bali); Pedinopleurus notabilis (Silvestri, 1946); Aenictoides gen. nov. derivata spec. nov. (Thailand). The intrageneric affiliations of Orphnebius Motschulsky, 1858 and the subgeneric concept currently in use are discussed, and new species (sub-)groups are proposed. The following synonymies are proposed: Deroleptus Bernhauer, 1915 = Megalocephalobius Bernhauer, 1929, syn. nov.; Orphnebius breviceps Cameron, 1946 = O. vorax Pace, 2000, syn. nov.; Orphnebius niger (Cameron, 1939) = O. turensis Pace, 2012, syn. nov.; Drusilla canaliculata (Fabricius, 1787) = Myrmedonia polyporina Gistel, 1857, syn. nov.; Drusilla erichsoni (Peyron, 1857) = Myrmedonia aptera Peyron, 1858, syn. nov.; Drusilla zyrasoides Dvořak, 1988 = D. truncatella Pace, 2004, syn. nov.; Amaurodera kraepelini Fauvel, 1905 = A. rougemonti Pace, 1987, syn. nov., = A. smetanai Pace, 1992, syn. nov., = A. yaoana Pace, 1992, syn. nov.; Tetrabothrus bicolor Cameron, 1939 = T. chinensis Pace, 2012, syn. nov.; Tetrabothrus puetzi Assing, 2009 = T. taiwanensis Pace, 2010, syn. nov.; Tetrabothrus clavatus Bernhauer, 1915 = T. quadricollis Cameron, 1950, syn. nov., = T. japonicus Nakane, 1991, syn. nov., = T. vietnamiculus Pace, 2013, syn. nov., = T. rubricollis Assing, 2015, syn. nov.; Zyras fratrumkadooriorum Pace, 1998 = Z. chumphonensis Pace, 2004, syn. nov.; Pedinopleurus notabilis (Silvestri, 1946) = P. setosicauda Dvořák, 1996, syn. nov. Lectotypes are designated for Deroleptus niger Cameron, 1939, Orphnebius falagrioides Bernhauer, 1929, Myrmedonia aptera Peyron, 1858, and Tetrabothrus indicus Cameron, 1939. Orphnebius krypticola Pace, 2007 is designated as the correct original spelling. Updated catalogues of Amaurodera Fauvel, 1905 and Tetrabothrus Bernhauer, 1915, as well as keys to the Orphnebius species of Laos and to the Amaurodera species of Thailand are provided. Additional records of 41 named and several unnamed species are reported, among them numerous new country and province records. The distributions of Amaurodera kraepelini and Tetrabothrus clavatus are mapped.

Keywords

Coleoptera, Staphylinidae, Aleocharinae, Lomechusini, Palaearctic region, Oriental region, taxonomy, new genus, new species, new synonymies, lectotype designations, keys to species, catalogues, additional records, distribution maps.

Nomenclatural Acts

derivata Assing, 2016 (Aenictoides), spec. n.
arunica Assing, 2016 (Amaurodera), spec. n.
dentata Assing, 2016 (Amaurodera), spec. n.
disparicollis Assing, 2016 (Amaurodera), spec. n.
fasciata Assing, 2016 (Amaurodera), spec. n.
gilvios Assing, 2016 (Amaurodera), spec. n.
kraepelini rougemonti Pace, 1987 (Amaurodera), syn. n. of Amaurodera kraepelini Fauvel, 1905
parvoculata Assing, 2016 (Amaurodera), spec. n.
reticulata Assing, 2016 (Amaurodera), spec. n.
smetanai Pace, 1992a (Amaurodera), syn. n. of Amaurodera kraepelini Fauvel, 1905
spinans Assing, 2016 (Amaurodera), spec. n.
varicollis Assing, 2016 (Amaurodera), spec. n.
yaoana Pace, 1992b (Amaurodera), syn. n. of Amaurodera kraepelini Fauvel, 1905
bifida Assing, 2016 (Drusilla), spec. n.
truncatella Pace, 2004 (Drusilla), syn. n. of Drusilla zyrasoides Dvorák, 1988
Aenictoides Assing, 2016 (Lomechusini), gen. n.
aptera Peyron, 1858 (Myrmedonia), syn. n. of Drusilla (Drusilla) erichsoni (Peyron, 1857)
polyporina Gistel, 1857 (Myrmedonia), syn. n. of Drusilla (Drusilla) canaliculata (Fabricius, 1787)
bicuspis Assing, 2016 (Orphnebius), spec. n.
biformis Assing, 2016 (Orphnebius), spec. n.
cernens Assing, 2016 (Orphnebius), spec. n.
dilatatus Assing, 2016 (Orphnebius), spec. n.
effeminatus Assing, 2016 (Orphnebius), spec. n.
extensus Assing, 2016 (Orphnebius), spec. n.
falagrioides Bernhauer, 1929 (Orphnebius), LT; described as Orphnebius (Megalocephalobius) Bernhauer, 1929
fodens Assing, 2016 (Orphnebius), spec. n.
fuscapicalis Assing, 2016 (Orphnebius), spec. n.
fuscicollis Assing, 2016 (Orphnebius), spec. n.
grandicollis Assing, 2016 (Orphnebius), spec. n.
integer Assing, 2016 (Orphnebius), spec. n.
latitibialis Assing, 2016 (Orphnebius), spec. n.
lunatus Assing, 2016 (Orphnebius), spec. n.
nigrapicalis Assing, 2016 (Orphnebius), spec. n.
reductus Assing, 2016 (Orphnebius), spec. n.
retunsus Assing, 2016 (Orphnebius), spec. n.
serratus Assing, 2016 (Orphnebius), spec. n.
spinans Assing, 2016 (Orphnebius), spec. n.
turensis Pace, 2012b (Orphnebius), syn. n. of Orphnebius (Deroleptus) niger (Cameron, 1939)
vorax Pace, 2000 (Orphnebius), syn. n. of Orphnebius breviceps Cameron, 1946
baccillatus Assing, 2016 (Orphnebius (Deroleptus)), spec. n.
biimpressus Assing, 2016 (Orphnebius (Deroleptus)), spec. n.
carinatus Assing, 2016 (Orphnebius (Deroleptus)), spec. n.
cultellatus Assing, 2016 (Orphnebius (Deroleptus)), spec. n.
discrepans Assing, 2016 (Orphnebius (Deroleptus)), spec. n.
dispar Assing, 2016 (Orphnebius (Deroleptus)), spec. n.
gracilior Assing, 2016 (Orphnebius (Deroleptus)), spec. n.
septemcuspis Assing, 2016 (Orphnebius (Deroleptus)), spec. n.
sexcarinatus Assing, 2016 (Orphnebius (Deroleptus)), spec. n.
spoliatus Assing, 2016 (Orphnebius (Deroleptus)), spec. n.
tortus Assing, 2016 (Orphnebius (Deroleptus)), spec. n.
triapicalis Assing, 2016 (Orphnebius (Deroleptus)), spec. n.
ulcerosus Assing, 2016 (Orphnebius (Deroleptus)), spec. n.
vates Assing, 2016 (Orphnebius (Deroleptus)), spec. n.
pectinata Assing, 2016 (Rabdotodrusilla), spec. n.
breviatus Assing, 2016 (Tetrabothrus), spec. n.
chinensis Pace, 2012a (Tetrabothrus), syn. n. of Tetrabothrus bicolor Cameron, 1939
collucatus Assing, 2016 (Tetrabothrus), spec. n.
japonicus Nakane, 1991 (Tetrabothrus), syn. n. of Tetrabothrus clavatus Bernhauer, 1915
nilgiricus Assing, 2016 (Tetrabothrus), spec. n.
punctiventris Assing, 2016 (Tetrabothrus), spec. n.
quadricollis Cameron, 1950 (Tetrabothrus), syn. n. of Tetrabothrus clavatus Bernhauer, 1915
rubricollis Assing, 2015 (Tetrabothrus), syn. n. of Tetrabothrus clavatus Bernhauer, 1915
sulawesicus Assing, 2016 (Tetrabothrus), spec. n.
taiwanensis Pace, 2010 (Tetrabothrus), syn. n. of Tetrabothrus puetzi Assing, 2009
vietnamiculus Pace, 2013 (Tetrabothrus), syn. n. of Tetrabothrus clavatus Bernhauer, 1915
chumphonensis Pace, 2004 (Zyras (Zyras)), syn. n. of Zyras (Zyras) fratrumkadooriorum Pace, 1998
porrectus Assing, 2016 (Zyras (Zyras)), spec. n.
wunderlei Assing, 2016 (Zyras (Zyras)), spec. n.