On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)
DOI:
https://doi.org/10.21248/contrib.entomol.67.2.213-246Abstract
Species of the nominal subgenus of the lomechusine genus Zyras Stephens, 1835 of the southeastern Oriental region are revised. Eighteen species are (re-)described and/or illustrated, six of them for the first time: Zyras (Zyras) latibasalis spec. nov. (Java, tentatively recorded also from Laos), Z. (Z.) rutrilobatus spec. nov. (Laos), Z. (Z.) parvilobatus spec. nov. (Laos), Z. (Z.) hirtiventris spec. nov. (Laos, Vietnam), Z. (Z.) hlavaci spec. nov. (Malaysia: Pahang, Selangor), Z. (Z.) lunatus spec. nov. (Malaysia: Selangor). Eight synonymies are proposed: Zyras bryanti Cameron, 1943 = Z. mortuorum Pace, 1990, syn. nov., = Z. paederinus Pace, 2008, syn. nov.; Zyras nigerrimus Cameron, 1943 = Z. bartolozzii Pace, 2003, syn. nov., = Z. alboterminalis Pace, 2008, syn. nov.; Z. granulipennis Cameron, 1930 = Z. pervariolosus Pace, 2008, syn. nov.; Z. matangensis Cameron, 1943 = Z. daiaccorum Pace, 2008, syn. nov.; Z. montanus (Bernhauer, 1915) = Z. variolatus Pace, 2003, syn. nov.; Z. preangeranus Cameron, 1939 = Z. quadriterminalis Pace, 2008, syn. nov. One species is excluded from Zyras: Myrmedonota modiglianii (Cameron, 1925), comb. nov. Lectotypes are designated for Zyras elegantulus Cameron, 1939, Z. semirufus Cameron, 1939, and Z. flavus Cameron, 1939. Additional records of nine named species are reported. A key to the species recorded from the southeastern Oriental region and an updated catalogue of the species of the Palaearctic and Oriental regions are provided. The subgenus is currently represented in the Palaearctic and Oriental regions by a total of 122 described species. Twenty-two species have been recorded from Sunda Islands. The countries with the greatest diversity are China (46 named species), India (20), Indonesia (19), Malaysia (15), Thailand (14), Nepal (12), Laos (11), and Vietnam (8).
Nomenclatural Acts
- Zyras (Zyras) latibasalis spec. nov. – urn:lsid:zoobank.org:act:35276B1A-6492-4F55-8836-91058719625C
- Zyras (Zyras) rutrilobatus spec. nov. – urn:lsid:zoobank.org:act:67BAD12F-50BC-4983-A8BC-363FA67F01F1
- Zyras (Zyras) parvilobatus spec. nov. – urn:lsid:zoobank.org:act:6DBB36A1-C12B-4FA1-B28E-55560D6645C6
- Zyras (Zyras) hirtiventris spec. nov. – urn:lsid:zoobank.org:act:73489C67-FB52-4B48-A2E1-BA1F1E55F45D
- Zyras (Zyras) hlavaci spec. nov. – urn:lsid:zoobank.org:act:F5734436-CE49-4064-8A52-BC7DF9A0EE67
- Zyras (Zyras) lunatus spec. nov. – urn:lsid:zoobank.org:act:ACD6BAAB-FE21-415D-903F-1D2CDEFE0607
References
<DIV CLASS="zitation"><SPAN CLASS="autor">Assing, V.</SPAN> 2015: On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Staphylininae). – Linzer Biologische Beiträge <B>47</B>(2): 1257-1282 - <A HREF="http://www.zobodat.at/pdf/LBB_0047_2_1257-1282.pdf" TARGET="_blank">http://www.zobodat.at/pdf/LBB_0047_2_1257-1282.pdf</A>.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Assing, V.</SPAN> 2016a: A revision of Zyras Stephens sensu strictu of China, Taiwan, and Hong Kong, with records and (re-)descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). – Stuttgarter Beiträge zur Naturkunde A, Neue Serie <B>9</B>: 87-175.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Assing, V.</SPAN> 2016b: On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae). – Contributions to Entomology <B>66</B>(1): 13-111 - DOI: <A HREF="https://doi.org/10.21248/contrib.entomol.66.1.013-111" TARGET="_blank">10.21248/contrib.entomol.66.1.013-111</A>.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Assing, V.</SPAN> 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). – Contributions to Entomology <B>67</B>(1): 117-192 - DOI: <A HREF="https://doi.org/10.21248/contrib.entomol.67.1.117-192" TARGET="_blank">10.21248/contrib.entomol.67.1.117-192</A>.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Bernhauer, M.</SPAN> 1914: Neue Staphyliniden der paläarktischen Fauna. – Coleopterologische Rundschau <B>3</B>(4): 65-68.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Bernhauer, M.</SPAN> 1915: Neue Staphyliniden der indo-malaiischen Fauna, insbesondere der Sunda-Insel Borneo (9. Beitrag). – Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien <B>65</B>: 134-158.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Bernhauer, M.</SPAN> 1933a: Neuheiten der chinesischen Staphylinidenfauna. – Wiener Entomologische Zeitung <B>50</B>: 25-48.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Bernhauer, M.</SPAN> 1933b: Neues aus der Staphylinidenfauna China´s. – Entomologisches Nachrichtenblatt <B>7</B>(2): 39-54.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Bernhauer, M.</SPAN> 1939: Zur Staphylinidenfauna von China u. Japan. – Entomologisches Nachrichtenblatt <B>12</B>(3-4): 145-158.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Cameron, M.</SPAN> 1925: Descriptions of new species of oriental Staphylinidae. – Annali del Museo Civico di Storia Naturale di Genova <B>52</B>: 34-49.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Cameron, M.</SPAN> 1930: Staphylinidae from British North Borneo, with descriptions of new species. – Journal of the Federated Malay States Museums <B>16</B>: 160-168.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Cameron, M.</SPAN> 1939a: Coleoptera: Staphylinidae. Volume IV, Parts I-II - <SPAN CLASS="autor">Sewell, R. B. S.</SPAN> (ed): The Fauna of British India, including Ceylon and Burma., London: 691 pp.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Cameron, M.</SPAN> 1939b: Fauna Javanica. The Staphylinidae collected by Mr. F. C. Drescher. Part III. – Tijdschrift voor Entomologie <B>82</B>: 1-29.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Cameron, M.</SPAN> 1943: New species of Staphylinidae (Col.) from Borneo. – Entomologist's Monthly Magazine <B>79</B>: 39-42, 139-143, 183.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Cameron, M.</SPAN> 1944: Descriptions of new Staphylinidae (Coleoptera). – Proceedings of the Royal Entomological Society of London (B) <B>13</B>: 11-15, 49-52, 104-108.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Champion, G. C.</SPAN> 1921: Some Indian Coleoptera (6). – The Entomologist´s Monthly Magazine <B>57</B>: 177-184, 201-206.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Champion, G. C.</SPAN> 1927: Some Indian Coleoptera (23). – The Entomologist´s Monthly Magazine <B>63</B>: 245-250.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Dvořák, M.</SPAN> 1981: Zwei neue ostasiatische Arten und nomenklatorische Bemerkungen zur Gattung Zyras (Coleoptera, Staphylinidae). – Acta Entomologica Bohemoslovaca <B>78</B>: 53-60.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Dvořák, M.</SPAN> 1984: Zur Kenntnis einiger myrmekophiler Staphylinidae (Coleoptera). – Acta Entomologica Bohemoslovaca <B>81</B>: 190-203.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Dvořák, M.</SPAN> 1996: Einige neue myrmekophile oder termitophile Arten des Tribus Zyrasini aus Südasien (Coleoptera, Staphylinidae, Aleocharinae). – Coleoptera Schwanfelder Coleopterologische Mitteilungen <B>23</B>: 1-15.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Fauvel, A.</SPAN> 1904: Staphylinides de l'Hindoustan et de la Birmanie. – Revue d'Entomologie <B>23</B>: 43-70.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Hlaváč, P.; Newton, A. F. & Maruyama, M.</SPAN> 2011: World catalogue of the species of the tribe Lomechusini (Staphylinidae: Aleocharinae). – Zootaxa <B>3075</B>: 1-151.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Kraatz, G.</SPAN> 1859: Die Staphylinen-Fauna von Ostindien, insbesondere der Insel Ceylan. – Archiv für Naturgeschichte <B>25</B>(1): 1-193.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Last, H. R.</SPAN> 1969: Changes in the nomenclature of the genus Zyras Stephens (Col., Staphylinidae). – Entomologist's Monthly Magazine <B>104 [1968]</B>: 279.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Last, H. R.</SPAN> 1982: New species of Zyras from Mongolia (Coleoptera: Staphylinidae). – Acta Zoologica Academiae Scientiarum Hungaricae <B>28</B>(1-2): 81-82.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Motschulsky, V.</SPAN> 1861: Essai d'un catalogue des insectes de l'ile Ceylan. 1-ière livraison. – Bulletin de la Société Impériale des Naturalistes de Moscou <B>34</B>(Première partie): 95-155.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1986a: Aleocharinae della Thailandia e della Birmania riportate da G. de Rougemont (Coleoptera, Staphylinidae). – Bollettino del Museo Civico di Storia Naturale di Verona <B>11 [1984]</B>: 427-468.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1986b: Aleocharinae dell'Asia sudorientale raccolte dal Dr. Osella (Coleoptera, Staphylinidae). – Bollettino del Museo Civico di Storia Naturale di Verona <B>11 [1984]</B>: 481-491.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1986c: Aleocharinae dell'Himalaya raccolte da Guillaume de Rougemont (Coleoptera Staphylinidae). – Bollettino del Museo Civico di Storia Naturale di Verona <B>12</B>: 165-191.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1987a: Staphylinidae dell'Himalaya nepalese. Aleocharinae raccolte dal Prof. Dr. J. Martens (Insecta: Coleoptera). – Courier des Forschungsinstitutes Senckenberg <B>93</B>: 383-441.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1987b: Aleocharinae dell'Asia sudorientale raccolte da G. de Rougemont (Coleoptera, Staphylinidae). – Bollettino del Museo Civico di Storia Naturale di Verona <B>13 [1986]</B>: 139-237.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1988: Aleocharinae dello Sri Lanka raccolte da Marc Tronquet (Coleoptera Staphylinidae). – Bollettino del Museo Civico di Storia Naturale di Verona <B>14 [1987]</B>: 315-338.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1990: Aleocharinae delle Filippine. 82º contributo alla conoscenza delle Aleocharinae. – <SPAN CLASS="autor">Berti, N.</SPAN> (ed.): Miscellanées sur les Staphylins. - Mémoires du Muséum National d'Histoire Naturelle (A, Zoologie) 147: 57-113.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1992: Aleocharinae nepalesi del Museo di Ginevra. Parte VI: Myrmedoniini (Coleoptera, Staphylinidae). – Revue Suisse de Zoologie <B>99</B>(1): 125-145.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1993: Aleocharinae della China (Coleoptera, Staphylinidae). – Bollettino del Museo Civico di Storia Naturale di Verona <B>17 [1990]</B>: 69-126.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1998: Aleocharinae della Cina: Parte IV (Coleoptera, Staphylinidae). – Revue Suisse de Zoologie <B>105</B>(4): 911-982.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1999: Aleocharinae di Hong Kong (Coleoptera, Staphylinidae). – Revue Suisse de Zoologie <B>106</B>(3): 663-689.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2001: Aleocharinae aus Vietnam in der Sammlung des Naturkundemuseums Erfurt (Coleoptera, Staphylinidae). – Veröffentlichungen Naturkundemuseum Erfurt <B>20</B>: 193-200.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2003: Aleocharinae della Penisola di Malacca (Malaysia) (Coleoptera, Staphylinidae). – Bollettino del Museo Regionale di Scienze Naturali, Torino <B>20</B>(1): 33-78.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2004: Aleocharinae di Thailandia, Cambogia, Laos e Malaysia (Coleoptera, Staphylinidae). – Belgian Journal of Entomology <B>6</B>: 243-302.</DIV>
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<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2006: Aleocharinae del Nepal al Naturkundemuseum di Erfurt (Insecta: Coleoptera: Staphylinidae). 343-408 – In: <SPAN CLASS="autor">Hartmann, M. & Weipert, J.</SPAN> (eds): Biodiversität und Naturausstattung im Himalaya. Volume II – Verein der Freunde und Förderer des Naturkundemuseums Erfurt e.V., Erfurt.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2008: Generi e specie della tribù Lomechusini del Borneo (Coleoptera, Staphylinidae). – Revue Suisse de Zoologie <B>115</B>(1): 107-156.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2010: Aleocharinae della regione Orientale al Museo di Genova (Coleoptera, Staphylinidae). – Annali del Museo Civico di Storia Naturale “G. Doria” <B>102</B>: 295-335.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2011: Aleocharinae from India gathered by Guillaume de Rougemont (Coleoptera, Staphylinidae). – Lavori della Società Veneziana di Scienze Naturali <B>36</B>: 23-40.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2012a: Biodiversità delle Aleocharinae della Cina: Lomechusini e Thamiaraeini (Coleoptera, Staphylinidae). – Beiträge zur Entomologie <B>62</B>(1): 77-102 - DOI: <A HREF="https://doi.org/10.21248/contrib.entomol.62.1.077-102" TARGET="_blank">10.21248/contrib.entomol.62.1.077-102</A>.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2012b: New data, new species, and two new genera of Aleocharinae from the Oriental Region (Insecta: Coleoptera: Staphylinidae). – Vernate <B>31</B>: 319-360.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2013: Nuovo contributo alla conoscenza delle Aleocharinae del Nepal (Insecta: Coleoptera: Staphylinidae). – Vernate <B>32</B>: 347-370.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2014: Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae). – Linzer Biologische Beiträge <B>46</B>(1): 727-794.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Scheerpeltz, O.</SPAN> 1965: Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Coleoptera Staphylinidae (except. Megalopsidiinae et Steninae). – Arkiv för Zoologie <B>17</B>(2): 93-371.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Schubert, K.</SPAN> 1908: Beitrag zur Staphylinidenfauna Ostindiens (West-Himalaya). (Col.) – Deutsche Entomologische Zeitschrift <B>1908</B>: 609-625.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Schülke, M. & Smetana, A.</SPAN> 2015: Staphylinidae.: 304-1134 – In: <SPAN CLASS="autor">Löbl, I. & Löbl, D.</SPAN> (eds): Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea – Staphylinoidea., Revised and updated Edition – Brill, Leiden – ISBN 9789004289925.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Assing, V.</SPAN> 2016a: A revision of Zyras Stephens sensu strictu of China, Taiwan, and Hong Kong, with records and (re-)descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). – Stuttgarter Beiträge zur Naturkunde A, Neue Serie <B>9</B>: 87-175.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Assing, V.</SPAN> 2016b: On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae). – Contributions to Entomology <B>66</B>(1): 13-111 - DOI: <A HREF="https://doi.org/10.21248/contrib.entomol.66.1.013-111" TARGET="_blank">10.21248/contrib.entomol.66.1.013-111</A>.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Assing, V.</SPAN> 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). – Contributions to Entomology <B>67</B>(1): 117-192 - DOI: <A HREF="https://doi.org/10.21248/contrib.entomol.67.1.117-192" TARGET="_blank">10.21248/contrib.entomol.67.1.117-192</A>.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Bernhauer, M.</SPAN> 1914: Neue Staphyliniden der paläarktischen Fauna. – Coleopterologische Rundschau <B>3</B>(4): 65-68.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Bernhauer, M.</SPAN> 1915: Neue Staphyliniden der indo-malaiischen Fauna, insbesondere der Sunda-Insel Borneo (9. Beitrag). – Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien <B>65</B>: 134-158.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Bernhauer, M.</SPAN> 1933a: Neuheiten der chinesischen Staphylinidenfauna. – Wiener Entomologische Zeitung <B>50</B>: 25-48.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Bernhauer, M.</SPAN> 1933b: Neues aus der Staphylinidenfauna China´s. – Entomologisches Nachrichtenblatt <B>7</B>(2): 39-54.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Bernhauer, M.</SPAN> 1939: Zur Staphylinidenfauna von China u. Japan. – Entomologisches Nachrichtenblatt <B>12</B>(3-4): 145-158.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Cameron, M.</SPAN> 1925: Descriptions of new species of oriental Staphylinidae. – Annali del Museo Civico di Storia Naturale di Genova <B>52</B>: 34-49.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Cameron, M.</SPAN> 1930: Staphylinidae from British North Borneo, with descriptions of new species. – Journal of the Federated Malay States Museums <B>16</B>: 160-168.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Cameron, M.</SPAN> 1939a: Coleoptera: Staphylinidae. Volume IV, Parts I-II - <SPAN CLASS="autor">Sewell, R. B. S.</SPAN> (ed): The Fauna of British India, including Ceylon and Burma., London: 691 pp.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Cameron, M.</SPAN> 1939b: Fauna Javanica. The Staphylinidae collected by Mr. F. C. Drescher. Part III. – Tijdschrift voor Entomologie <B>82</B>: 1-29.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Cameron, M.</SPAN> 1943: New species of Staphylinidae (Col.) from Borneo. – Entomologist's Monthly Magazine <B>79</B>: 39-42, 139-143, 183.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Cameron, M.</SPAN> 1944: Descriptions of new Staphylinidae (Coleoptera). – Proceedings of the Royal Entomological Society of London (B) <B>13</B>: 11-15, 49-52, 104-108.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Champion, G. C.</SPAN> 1921: Some Indian Coleoptera (6). – The Entomologist´s Monthly Magazine <B>57</B>: 177-184, 201-206.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Champion, G. C.</SPAN> 1927: Some Indian Coleoptera (23). – The Entomologist´s Monthly Magazine <B>63</B>: 245-250.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Dvořák, M.</SPAN> 1981: Zwei neue ostasiatische Arten und nomenklatorische Bemerkungen zur Gattung Zyras (Coleoptera, Staphylinidae). – Acta Entomologica Bohemoslovaca <B>78</B>: 53-60.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Dvořák, M.</SPAN> 1984: Zur Kenntnis einiger myrmekophiler Staphylinidae (Coleoptera). – Acta Entomologica Bohemoslovaca <B>81</B>: 190-203.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Dvořák, M.</SPAN> 1996: Einige neue myrmekophile oder termitophile Arten des Tribus Zyrasini aus Südasien (Coleoptera, Staphylinidae, Aleocharinae). – Coleoptera Schwanfelder Coleopterologische Mitteilungen <B>23</B>: 1-15.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Fauvel, A.</SPAN> 1904: Staphylinides de l'Hindoustan et de la Birmanie. – Revue d'Entomologie <B>23</B>: 43-70.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Hlaváč, P.; Newton, A. F. & Maruyama, M.</SPAN> 2011: World catalogue of the species of the tribe Lomechusini (Staphylinidae: Aleocharinae). – Zootaxa <B>3075</B>: 1-151.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Kraatz, G.</SPAN> 1859: Die Staphylinen-Fauna von Ostindien, insbesondere der Insel Ceylan. – Archiv für Naturgeschichte <B>25</B>(1): 1-193.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Last, H. R.</SPAN> 1969: Changes in the nomenclature of the genus Zyras Stephens (Col., Staphylinidae). – Entomologist's Monthly Magazine <B>104 [1968]</B>: 279.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Last, H. R.</SPAN> 1982: New species of Zyras from Mongolia (Coleoptera: Staphylinidae). – Acta Zoologica Academiae Scientiarum Hungaricae <B>28</B>(1-2): 81-82.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Motschulsky, V.</SPAN> 1861: Essai d'un catalogue des insectes de l'ile Ceylan. 1-ière livraison. – Bulletin de la Société Impériale des Naturalistes de Moscou <B>34</B>(Première partie): 95-155.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1986a: Aleocharinae della Thailandia e della Birmania riportate da G. de Rougemont (Coleoptera, Staphylinidae). – Bollettino del Museo Civico di Storia Naturale di Verona <B>11 [1984]</B>: 427-468.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1986b: Aleocharinae dell'Asia sudorientale raccolte dal Dr. Osella (Coleoptera, Staphylinidae). – Bollettino del Museo Civico di Storia Naturale di Verona <B>11 [1984]</B>: 481-491.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1986c: Aleocharinae dell'Himalaya raccolte da Guillaume de Rougemont (Coleoptera Staphylinidae). – Bollettino del Museo Civico di Storia Naturale di Verona <B>12</B>: 165-191.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1987a: Staphylinidae dell'Himalaya nepalese. Aleocharinae raccolte dal Prof. Dr. J. Martens (Insecta: Coleoptera). – Courier des Forschungsinstitutes Senckenberg <B>93</B>: 383-441.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1987b: Aleocharinae dell'Asia sudorientale raccolte da G. de Rougemont (Coleoptera, Staphylinidae). – Bollettino del Museo Civico di Storia Naturale di Verona <B>13 [1986]</B>: 139-237.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1988: Aleocharinae dello Sri Lanka raccolte da Marc Tronquet (Coleoptera Staphylinidae). – Bollettino del Museo Civico di Storia Naturale di Verona <B>14 [1987]</B>: 315-338.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1990: Aleocharinae delle Filippine. 82º contributo alla conoscenza delle Aleocharinae. – <SPAN CLASS="autor">Berti, N.</SPAN> (ed.): Miscellanées sur les Staphylins. - Mémoires du Muséum National d'Histoire Naturelle (A, Zoologie) 147: 57-113.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1992: Aleocharinae nepalesi del Museo di Ginevra. Parte VI: Myrmedoniini (Coleoptera, Staphylinidae). – Revue Suisse de Zoologie <B>99</B>(1): 125-145.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1993: Aleocharinae della China (Coleoptera, Staphylinidae). – Bollettino del Museo Civico di Storia Naturale di Verona <B>17 [1990]</B>: 69-126.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1998: Aleocharinae della Cina: Parte IV (Coleoptera, Staphylinidae). – Revue Suisse de Zoologie <B>105</B>(4): 911-982.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 1999: Aleocharinae di Hong Kong (Coleoptera, Staphylinidae). – Revue Suisse de Zoologie <B>106</B>(3): 663-689.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2001: Aleocharinae aus Vietnam in der Sammlung des Naturkundemuseums Erfurt (Coleoptera, Staphylinidae). – Veröffentlichungen Naturkundemuseum Erfurt <B>20</B>: 193-200.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2003: Aleocharinae della Penisola di Malacca (Malaysia) (Coleoptera, Staphylinidae). – Bollettino del Museo Regionale di Scienze Naturali, Torino <B>20</B>(1): 33-78.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2004: Aleocharinae di Thailandia, Cambogia, Laos e Malaysia (Coleoptera, Staphylinidae). – Belgian Journal of Entomology <B>6</B>: 243-302.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2005: Nuovo contributo alla conoscenza delle Aleocharinae dell'India (Coleoptera, Staphylinidae). – Nouvelle Revue d'Entomologie (N.S.) <B>22</B>(2): 141-153.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2006: Aleocharinae del Nepal al Naturkundemuseum di Erfurt (Insecta: Coleoptera: Staphylinidae). 343-408 – In: <SPAN CLASS="autor">Hartmann, M. & Weipert, J.</SPAN> (eds): Biodiversität und Naturausstattung im Himalaya. Volume II – Verein der Freunde und Förderer des Naturkundemuseums Erfurt e.V., Erfurt.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2008: Generi e specie della tribù Lomechusini del Borneo (Coleoptera, Staphylinidae). – Revue Suisse de Zoologie <B>115</B>(1): 107-156.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2010: Aleocharinae della regione Orientale al Museo di Genova (Coleoptera, Staphylinidae). – Annali del Museo Civico di Storia Naturale “G. Doria” <B>102</B>: 295-335.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2011: Aleocharinae from India gathered by Guillaume de Rougemont (Coleoptera, Staphylinidae). – Lavori della Società Veneziana di Scienze Naturali <B>36</B>: 23-40.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2012a: Biodiversità delle Aleocharinae della Cina: Lomechusini e Thamiaraeini (Coleoptera, Staphylinidae). – Beiträge zur Entomologie <B>62</B>(1): 77-102 - DOI: <A HREF="https://doi.org/10.21248/contrib.entomol.62.1.077-102" TARGET="_blank">10.21248/contrib.entomol.62.1.077-102</A>.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2012b: New data, new species, and two new genera of Aleocharinae from the Oriental Region (Insecta: Coleoptera: Staphylinidae). – Vernate <B>31</B>: 319-360.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2013: Nuovo contributo alla conoscenza delle Aleocharinae del Nepal (Insecta: Coleoptera: Staphylinidae). – Vernate <B>32</B>: 347-370.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Pace, R.</SPAN> 2014: Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae). – Linzer Biologische Beiträge <B>46</B>(1): 727-794.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Scheerpeltz, O.</SPAN> 1965: Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Coleoptera Staphylinidae (except. Megalopsidiinae et Steninae). – Arkiv för Zoologie <B>17</B>(2): 93-371.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Schubert, K.</SPAN> 1908: Beitrag zur Staphylinidenfauna Ostindiens (West-Himalaya). (Col.) – Deutsche Entomologische Zeitschrift <B>1908</B>: 609-625.</DIV>
<DIV CLASS="zitation"><SPAN CLASS="autor">Schülke, M. & Smetana, A.</SPAN> 2015: Staphylinidae.: 304-1134 – In: <SPAN CLASS="autor">Löbl, I. & Löbl, D.</SPAN> (eds): Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea – Staphylinoidea., Revised and updated Edition – Brill, Leiden – ISBN 9789004289925.</DIV>
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2017-12-08
How to Cite
Assing, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini) - Contributions to Entomology = Beiträge Zur Entomologie 67(2): 213–246 - doi: 10.21248/contrib.entomol.67.2.213-246
